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A friend of mine who's not with us anymore :( had a MK5 ZS with a puma lump in it. It kept up with my C2 vts I had at the time.
And my C2 was not even close to my clio.
ECU (brain of the vehicle) - basically finely tunes the brain of the vehicle which controls the engine if you like for more specific values, whether that be reducing flat spots, creating more power or better mpg etc.
It's a more 'honed-optimised' brain for your car.